{"id":181,"date":"2017-09-13T20:20:55","date_gmt":"2017-09-13T20:20:55","guid":{"rendered":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/chapter\/25-homo-species-indeterminate\/"},"modified":"2017-09-13T20:20:55","modified_gmt":"2017-09-13T20:20:55","slug":"25-homo-species-indeterminate","status":"publish","type":"chapter","link":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/chapter\/25-homo-species-indeterminate\/","title":{"raw":"25. Homo species indeterminate","rendered":"25. Homo species indeterminate"},"content":{"raw":"<h1><i>Homo<\/i><b> species indeterminate (2.8 mya)<\/b><\/h1>\nA possible new species of Early <i>Homo<\/i> was recently discovered. Villmoare et al. (2015) reported on what they are calling <i>Homo<\/i> species indet. (\u201cindeterminate\u201d). The half mandible (LD 350-1) was discovered by Chalachew Seyoum in 2013 at the Ledi-Geraru site in the Afar region of Ethiopia. If indeed the jaw is part of the <i>Homo<\/i> lineage, it extends the origin of our genus to 2.8 mya, 400 kya older than the oldest <i>H. habilis<\/i> specimen, i.e. AL 666-1 from the Hadar site in Ethiopia where Lucy was discovered. Villmoare et al. (2015) believe that the specimen represents a transitional form between <i>Au. afarensis<\/i> and <i>Homo habilis<\/i>. Support for their supposition includes the combination of australopith and <i>Homo<\/i>-like characteristics seen in the mandible and teeth, as well as the fact that it is from the same area as <i>Au. afarensis<\/i> that is known to have survived there until 3 mya. They report on a fragmentary mandible from the Koobi Fora site in Kenya (KNM-ER 5431) that exhibits a similar combination of australopith and <i>Homo<\/i> morphology (see Villmoare et al. 2015 for references).\n\nThe anterior of the mandible exhibits the most primitive characteristics, along with some <i>Homo<\/i>-like characteristics. <i>Homo<\/i> characteristics include narrower molars with more derived dimensions and cusp morphology, and the canine\/first premolar configuration is less ape- or australopith-like, as evidenced by wear patterns. They conclude that deviation in the teeth and jaws, from the australopith condition, occurred early in the <i>Homo<\/i> lineage.\n\nDiMaggio et al. (2015) reported on the paleoenvironment of the site. The area was a mosaic environment consisting of open grass- and scrubland, gallery forest, and lakes and\/or rivers. They see a faunal turnover 2.8\u20132.6 mya in the strata, in accordance with the area becoming more open and likely arid with an increase in grazers. They cite evidence for climatic and vegetation shifts resulting from \u201crifting processes and extensive volcanism [that] altered the architecture of sedimentary basins\u201d (see DiMaggio et al., p. 1, for references). Their work fills an important gap in the paleoenvironmental history of the region.\n\nThe fact that <i>Au. garhi<\/i> is younger and their teeth are more robust leads the researchers to conclude parsimoniously that <i>Au. garhi<\/i> is not part of our ancestry, as earlier and later forms of <i>Homo<\/i> had more gracile teeth and jaws. The same would also hold true for <i>Au. sediba<\/i> due to their much later appearance in the fossil record at ~2 mya. However, Hawks et al. (2015) argue that the <b>osteometrics<\/b> (bone measurements) and anatomical features do not support firm taxonomic placement in genus:\u00a0<i>Homo<\/i>. They believe that they have ample evidence showing that <i>Au. sediba<\/i> is closely related to early species of <i>Homo<\/i> (<a href=\"..\/australopithecus-sediba\/\">see Chapter 21<\/a>).","rendered":"<h1><i>Homo<\/i><b> species indeterminate (2.8 mya)<\/b><\/h1>\n<p>A possible new species of Early <i>Homo<\/i> was recently discovered. Villmoare et al. (2015) reported on what they are calling <i>Homo<\/i> species indet. (\u201cindeterminate\u201d). The half mandible (LD 350-1) was discovered by Chalachew Seyoum in 2013 at the Ledi-Geraru site in the Afar region of Ethiopia. If indeed the jaw is part of the <i>Homo<\/i> lineage, it extends the origin of our genus to 2.8 mya, 400 kya older than the oldest <i>H. habilis<\/i> specimen, i.e. AL 666-1 from the Hadar site in Ethiopia where Lucy was discovered. Villmoare et al. (2015) believe that the specimen represents a transitional form between <i>Au. afarensis<\/i> and <i>Homo habilis<\/i>. Support for their supposition includes the combination of australopith and <i>Homo<\/i>-like characteristics seen in the mandible and teeth, as well as the fact that it is from the same area as <i>Au. afarensis<\/i> that is known to have survived there until 3 mya. They report on a fragmentary mandible from the Koobi Fora site in Kenya (KNM-ER 5431) that exhibits a similar combination of australopith and <i>Homo<\/i> morphology (see Villmoare et al. 2015 for references).<\/p>\n<p>The anterior of the mandible exhibits the most primitive characteristics, along with some <i>Homo<\/i>-like characteristics. <i>Homo<\/i> characteristics include narrower molars with more derived dimensions and cusp morphology, and the canine\/first premolar configuration is less ape- or australopith-like, as evidenced by wear patterns. They conclude that deviation in the teeth and jaws, from the australopith condition, occurred early in the <i>Homo<\/i> lineage.<\/p>\n<p>DiMaggio et al. (2015) reported on the paleoenvironment of the site. The area was a mosaic environment consisting of open grass- and scrubland, gallery forest, and lakes and\/or rivers. They see a faunal turnover 2.8\u20132.6 mya in the strata, in accordance with the area becoming more open and likely arid with an increase in grazers. They cite evidence for climatic and vegetation shifts resulting from \u201crifting processes and extensive volcanism [that] altered the architecture of sedimentary basins\u201d (see DiMaggio et al., p. 1, for references). Their work fills an important gap in the paleoenvironmental history of the region.<\/p>\n<p>The fact that <i>Au. garhi<\/i> is younger and their teeth are more robust leads the researchers to conclude parsimoniously that <i>Au. garhi<\/i> is not part of our ancestry, as earlier and later forms of <i>Homo<\/i> had more gracile teeth and jaws. The same would also hold true for <i>Au. sediba<\/i> due to their much later appearance in the fossil record at ~2 mya. However, Hawks et al. (2015) argue that the <b>osteometrics<\/b> (bone measurements) and anatomical features do not support firm taxonomic placement in genus:\u00a0<i>Homo<\/i>. They believe that they have ample evidence showing that <i>Au. sediba<\/i> is closely related to early species of <i>Homo<\/i> (<a href=\"..\/australopithecus-sediba\/\">see Chapter 21<\/a>).<\/p>\n\n\t\t\t <section class=\"citations-section\" role=\"contentinfo\">\n\t\t\t <h3>Candela Citations<\/h3>\n\t\t\t\t\t <div>\n\t\t\t\t\t\t <div id=\"citation-list-181\">\n\t\t\t\t\t\t\t <div class=\"licensing\"><div class=\"license-attribution-dropdown-subheading\">CC licensed content, Shared previously<\/div><ul class=\"citation-list\"><li>The History of our Tribe: Hominini. <strong>Authored by<\/strong>: Barbara Welker. <strong>Provided by<\/strong>: SUNY Geneseo. <strong>Located at<\/strong>: <a target=\"_blank\" href=\"https:\/\/milnepublishing.geneseo.edu\/the-history-of-our-tribe-hominini\/\">https:\/\/milnepublishing.geneseo.edu\/the-history-of-our-tribe-hominini\/<\/a>. <strong>Project<\/strong>: Open SUNY Textbooks. <strong>License<\/strong>: <em><a target=\"_blank\" rel=\"license\" href=\"https:\/\/creativecommons.org\/licenses\/by-nc-sa\/4.0\/\">CC BY-NC-SA: Attribution-NonCommercial-ShareAlike<\/a><\/em><\/li><\/ul><\/div>\n\t\t\t\t\t\t <\/div>\n\t\t\t\t\t <\/div>\n\t\t\t <\/section>","protected":false},"author":62,"menu_order":11,"template":"","meta":{"_candela_citation":"[{\"type\":\"cc\",\"description\":\"The History of our Tribe: Hominini\",\"author\":\"Barbara Welker\",\"organization\":\"SUNY Geneseo\",\"url\":\"https:\/\/milnepublishing.geneseo.edu\/the-history-of-our-tribe-hominini\/\",\"project\":\"Open SUNY Textbooks\",\"license\":\"cc-by-nc-sa\",\"license_terms\":\"\"}]","CANDELA_OUTCOMES_GUID":"","pb_show_title":"on","pb_short_title":"","pb_subtitle":"","pb_authors":[],"pb_section_license":""},"chapter-type":[],"contributor":[],"license":[],"class_list":["post-181","chapter","type-chapter","status-publish","hentry"],"part":144,"_links":{"self":[{"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/pressbooks\/v2\/chapters\/181","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/pressbooks\/v2\/chapters"}],"about":[{"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/wp\/v2\/types\/chapter"}],"author":[{"embeddable":true,"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/wp\/v2\/users\/62"}],"version-history":[{"count":0,"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/pressbooks\/v2\/chapters\/181\/revisions"}],"part":[{"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/pressbooks\/v2\/parts\/144"}],"metadata":[{"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/pressbooks\/v2\/chapters\/181\/metadata\/"}],"wp:attachment":[{"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/wp\/v2\/media?parent=181"}],"wp:term":[{"taxonomy":"chapter-type","embeddable":true,"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/pressbooks\/v2\/chapter-type?post=181"},{"taxonomy":"contributor","embeddable":true,"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/wp\/v2\/contributor?post=181"},{"taxonomy":"license","embeddable":true,"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/wp\/v2\/license?post=181"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}