{"id":216,"date":"2017-09-13T20:22:08","date_gmt":"2017-09-13T20:22:08","guid":{"rendered":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/chapter\/31-homo-antecessor\/"},"modified":"2017-10-03T16:22:58","modified_gmt":"2017-10-03T16:22:58","slug":"31-homo-antecessor","status":"publish","type":"chapter","link":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/chapter\/31-homo-antecessor\/","title":{"raw":"31. Homo antecessor","rendered":"31. Homo antecessor"},"content":{"raw":"<div class=\"thirtyone\">\r\n<h1><i>Homo antecessor<\/i><b> (1.2 mya)<\/b><\/h1>\r\n<h2>(\u201csame\u201d \/ \u201cpredecessor\u201d or \u201cpioneer\u201d)<\/h2>\r\n[caption id=\"attachment_313\" align=\"aligncenter\" width=\"288\"]<img class=\"wp-image-313 size-medium\" src=\"https:\/\/s3-us-west-2.amazonaws.com\/courses-images\/wp-content\/uploads\/sites\/2461\/2017\/09\/13202152\/image18-288x300.jpg\" alt=\"8.18\" width=\"288\" height=\"300\" \/> <em>Figure 31.1<\/em> Incomplete Homo antecessor skull from Gran Dolina, Spain.\u00a0\u201c<a href=\"https:\/\/commons.wikimedia.org\/wiki\/File:Homo_antecessor.jpg\">Homo antecessor<\/a>\u201d by Jos\u00e9-Manuel Benito is in the public domain.[\/caption]\r\n\r\n<div class=\"textbox examples\">\r\n<h3><b>SITES<\/b><\/h3>\r\nGran Dolina and Sima del Elefante, Spain\r\n<h3><b>PEOPLE<\/b><\/h3>\r\n<a href=\"http:\/\/en.wikipedia.org\/wiki\/Eudald_Carbonell\">Eudald Carbonell<\/a>, <a href=\"http:\/\/en.wikipedia.org\/wiki\/Juan_Luis_Arsuaga\">Juan Luis Arsuaga<\/a>, and J. M. Berm\u00fadez de Castro\r\n\r\n<\/div>\r\n<h2>INTRODUCTION<\/h2>\r\nUntil the recent discovery of hominin fossils dating to over 1.2 mya in the Atapuerca Mountains of Spain, the earliest known fossil material from the area was no more than 800 kya. The 800 kya material was assigned the taxonomic designation of <i>Homo antecessor<\/i>, or \u201cPioneer Man,\u201d as the first hominins to have ranged into Western Europe.\r\n<h2><b>PHYLOGENY<\/b><\/h2>\r\nSince its discovery, researchers have speculated that <i>Homo antecessor<\/i> was descended from a more derived form of <i>H. ergaster<\/i> that may have crossed the Mediterranean Sea from North Africa during a period of lowered sea levels. Now that the geological clock has been pushed back 300 kya, it is thought that they may have traveled through the Levant (i.e. the Eastern Mediterranean region). What is not known is whether the evolutionary event took place in Europe or North Africa. As far as descendant species, <i>Homo antecessor <\/i>is one contender for the ancestor of <i>Homo heidelbergensis<\/i> that in turn is thought to have given rise to neandertals and modern humans.\r\n\r\n[caption id=\"attachment_314\" align=\"aligncenter\" width=\"715\"]<img class=\"wp-image-314 size-full\" src=\"https:\/\/s3-us-west-2.amazonaws.com\/courses-images\/wp-content\/uploads\/sites\/2461\/2017\/09\/13202154\/H.-antecessor-e1487005782532.png\" alt=\"image\" width=\"715\" height=\"770\" \/> <em>Figure 31.2\u00a0<\/em>Homo antecessor by Keenan Taylor.[\/caption]\r\n<h2><b>DISCOVERY AND GEOGRAPHIC RANGE<\/b><\/h2>\r\nThe species was discovered by <a href=\"http:\/\/en.wikipedia.org\/wiki\/Eudald_Carbonell\">Eudald Carbonell<\/a>, <a href=\"http:\/\/en.wikipedia.org\/wiki\/Juan_Luis_Arsuaga\">Juan Luis Arsuaga<\/a>, and J. M. Berm\u00fadez de Castro. The first site to yield fossil material was the Gran Dolina site (see Figure 31.4) in the Atapuerca mountain range in Spain. Dated to 900 kya, the site has yielded both <i>H. antecessor<\/i> and <i>H. heidelbergensis<\/i> material. Eighty fossils from six individuals\u00a0were recovered. It is of interest that much of the fossil material is immature and exhibits cut marks that may indicate that they were victims of cannibalism.\r\n\r\n[caption id=\"attachment_266\" align=\"aligncenter\" width=\"750\"]<img class=\"wp-image-266\" src=\"https:\/\/s3-us-west-2.amazonaws.com\/courses-images\/wp-content\/uploads\/sites\/2461\/2017\/09\/13202158\/Mountain-Overlook-1024x523.png\" alt=\"image\" width=\"750\" height=\"383\" \/> <em>Figure 31.3\u00a0<\/em>\"Homo antecessor moves to Spain\" by Keenan Taylor.[\/caption]\r\n\r\nThe earliest <i>H. antecessor<\/i> site is Sima del Elefante (\u201cPit of the Elephant\u201d). Dated to 1.2 mya, only a handful of fossils have been recovered from the site.\r\n\r\nIn addition to other sites in Spain, tools and footprints have been found in Happisburgh, England and Ceprano, Italy (also known as <i>Homo cepranensis<\/i>, some consider it more closely related to <i>H. heidelbergensis<\/i>).\r\n\r\n[caption id=\"attachment_314\" align=\"aligncenter\" width=\"1432\"]<img class=\"wp-image-314 size-full\" src=\"https:\/\/s3-us-west-2.amazonaws.com\/courses-images\/wp-content\/uploads\/sites\/2461\/2017\/09\/13202200\/image19-1.jpg\" alt=\"8.19\" width=\"1432\" height=\"522\" \/> <em>Figure 31.4\u00a0<\/em>Gran Dolina site.\u00a0\u201c<a href=\"https:\/\/commons.wikimedia.org\/wiki\/File:Gran_Dolina_2012_(Sierra_de_Atapuerca).jpg\">Gran Dolina 2012 (Sierra de Atapuerca)<\/a>\u201d by Mario Modesto Mata is licensed under <a href=\"https:\/\/creativecommons.org\/licenses\/by-sa\/2.0\/deed.en\">CC BY-SA 2.0<\/a>.[\/caption]\r\n<h2><b>PHYSICAL CHARACTERISTICS<\/b><\/h2>\r\nThe 800 kya <i>H. antecessor<\/i> material was more derived than earlier <i>erectus<\/i> forms, possessing a larger brain (1,000\u20131,150 cc) and a more modern skull and face. They had low foreheads and possessed an <b>occipital bun<\/b> (a bun or chignon is the hairstyle wherein one winds long hair into a doughnut shape on the top or back of the head), as did neandertals (see neandertal features in Figure 31.5). It has been suggested that the purpose of that posterior protrusion is to balance the weight of the anterior portion of the skull and face. Based upon cranial anatomy, they are thought to have been capable of detecting the same range of sounds as modern humans and were possibly right-handed. They were almost as tall as modern humans at 5.5\u20136' (1.6\u20131.8 m), with males weighing ~200 lb (90\u00a0kg).\r\n\r\n[caption id=\"attachment_315\" align=\"aligncenter\" width=\"896\"]<img class=\"wp-image-315 size-full\" src=\"https:\/\/s3-us-west-2.amazonaws.com\/courses-images\/wp-content\/uploads\/sites\/2461\/2017\/09\/13202203\/image20-1.jpg\" alt=\"8.20\" width=\"896\" height=\"430\" \/> <em>Figure 31.5\u00a0<\/em>Neandertal cranial characteristics.\u00a0\u201c<a href=\"https:\/\/commons.wikimedia.org\/wiki\/File:Neanderthal_cranial_anatomy.jpg\">Neanderthal cranial anatomy<\/a>\u201d by Jason Potter is licensed under <a href=\"https:\/\/creativecommons.org\/licenses\/by-sa\/2.5\/deed.en\">CC BY-SA 2.5<\/a>.[\/caption]\r\n\r\n<\/div>\r\n<div class=\"thirtyone\">\r\n<div class=\"textbox key-takeaways\">\r\n<h3>Review of Derived Characteristics<\/h3>\r\n<ul>\r\n \t<li>Increased encephalization.<\/li>\r\n \t<li>Occipital bun.<\/li>\r\n \t<li>Tall like Turkana Boy.<\/li>\r\n<\/ul>\r\n<\/div>\r\n<\/div>","rendered":"<div class=\"thirtyone\">\n<h1><i>Homo antecessor<\/i><b> (1.2 mya)<\/b><\/h1>\n<h2>(\u201csame\u201d \/ \u201cpredecessor\u201d or \u201cpioneer\u201d)<\/h2>\n<div id=\"attachment_313\" style=\"width: 298px\" class=\"wp-caption aligncenter\"><img loading=\"lazy\" decoding=\"async\" aria-describedby=\"caption-attachment-313\" class=\"wp-image-313 size-medium\" src=\"https:\/\/s3-us-west-2.amazonaws.com\/courses-images\/wp-content\/uploads\/sites\/2461\/2017\/09\/13202152\/image18-288x300.jpg\" alt=\"8.18\" width=\"288\" height=\"300\" \/><\/p>\n<p id=\"caption-attachment-313\" class=\"wp-caption-text\"><em>Figure 31.1<\/em> Incomplete Homo antecessor skull from Gran Dolina, Spain.\u00a0\u201c<a href=\"https:\/\/commons.wikimedia.org\/wiki\/File:Homo_antecessor.jpg\">Homo antecessor<\/a>\u201d by Jos\u00e9-Manuel Benito is in the public domain.<\/p>\n<\/div>\n<div class=\"textbox examples\">\n<h3><b>SITES<\/b><\/h3>\n<p>Gran Dolina and Sima del Elefante, Spain<\/p>\n<h3><b>PEOPLE<\/b><\/h3>\n<p><a href=\"http:\/\/en.wikipedia.org\/wiki\/Eudald_Carbonell\">Eudald Carbonell<\/a>, <a href=\"http:\/\/en.wikipedia.org\/wiki\/Juan_Luis_Arsuaga\">Juan Luis Arsuaga<\/a>, and J. M. Berm\u00fadez de Castro<\/p>\n<\/div>\n<h2>INTRODUCTION<\/h2>\n<p>Until the recent discovery of hominin fossils dating to over 1.2 mya in the Atapuerca Mountains of Spain, the earliest known fossil material from the area was no more than 800 kya. The 800 kya material was assigned the taxonomic designation of <i>Homo antecessor<\/i>, or \u201cPioneer Man,\u201d as the first hominins to have ranged into Western Europe.<\/p>\n<h2><b>PHYLOGENY<\/b><\/h2>\n<p>Since its discovery, researchers have speculated that <i>Homo antecessor<\/i> was descended from a more derived form of <i>H. ergaster<\/i> that may have crossed the Mediterranean Sea from North Africa during a period of lowered sea levels. Now that the geological clock has been pushed back 300 kya, it is thought that they may have traveled through the Levant (i.e. the Eastern Mediterranean region). What is not known is whether the evolutionary event took place in Europe or North Africa. As far as descendant species, <i>Homo antecessor <\/i>is one contender for the ancestor of <i>Homo heidelbergensis<\/i> that in turn is thought to have given rise to neandertals and modern humans.<\/p>\n<div id=\"attachment_314\" style=\"width: 725px\" class=\"wp-caption aligncenter\"><img loading=\"lazy\" decoding=\"async\" aria-describedby=\"caption-attachment-314\" class=\"wp-image-314 size-full\" src=\"https:\/\/s3-us-west-2.amazonaws.com\/courses-images\/wp-content\/uploads\/sites\/2461\/2017\/09\/13202154\/H.-antecessor-e1487005782532.png\" alt=\"image\" width=\"715\" height=\"770\" \/><\/p>\n<p id=\"caption-attachment-314\" class=\"wp-caption-text\"><em>Figure 31.2\u00a0<\/em>Homo antecessor by Keenan Taylor.<\/p>\n<\/div>\n<h2><b>DISCOVERY AND GEOGRAPHIC RANGE<\/b><\/h2>\n<p>The species was discovered by <a href=\"http:\/\/en.wikipedia.org\/wiki\/Eudald_Carbonell\">Eudald Carbonell<\/a>, <a href=\"http:\/\/en.wikipedia.org\/wiki\/Juan_Luis_Arsuaga\">Juan Luis Arsuaga<\/a>, and J. M. Berm\u00fadez de Castro. The first site to yield fossil material was the Gran Dolina site (see Figure 31.4) in the Atapuerca mountain range in Spain. Dated to 900 kya, the site has yielded both <i>H. antecessor<\/i> and <i>H. heidelbergensis<\/i> material. Eighty fossils from six individuals\u00a0were recovered. It is of interest that much of the fossil material is immature and exhibits cut marks that may indicate that they were victims of cannibalism.<\/p>\n<div id=\"attachment_266\" style=\"width: 760px\" class=\"wp-caption aligncenter\"><img loading=\"lazy\" decoding=\"async\" aria-describedby=\"caption-attachment-266\" class=\"wp-image-266\" src=\"https:\/\/s3-us-west-2.amazonaws.com\/courses-images\/wp-content\/uploads\/sites\/2461\/2017\/09\/13202158\/Mountain-Overlook-1024x523.png\" alt=\"image\" width=\"750\" height=\"383\" \/><\/p>\n<p id=\"caption-attachment-266\" class=\"wp-caption-text\"><em>Figure 31.3\u00a0<\/em>&#8220;Homo antecessor moves to Spain&#8221; by Keenan Taylor.<\/p>\n<\/div>\n<p>The earliest <i>H. antecessor<\/i> site is Sima del Elefante (\u201cPit of the Elephant\u201d). Dated to 1.2 mya, only a handful of fossils have been recovered from the site.<\/p>\n<p>In addition to other sites in Spain, tools and footprints have been found in Happisburgh, England and Ceprano, Italy (also known as <i>Homo cepranensis<\/i>, some consider it more closely related to <i>H. heidelbergensis<\/i>).<\/p>\n<div id=\"attachment_314\" style=\"width: 1442px\" class=\"wp-caption aligncenter\"><img loading=\"lazy\" decoding=\"async\" aria-describedby=\"caption-attachment-314\" class=\"wp-image-314 size-full\" src=\"https:\/\/s3-us-west-2.amazonaws.com\/courses-images\/wp-content\/uploads\/sites\/2461\/2017\/09\/13202200\/image19-1.jpg\" alt=\"8.19\" width=\"1432\" height=\"522\" \/><\/p>\n<p id=\"caption-attachment-314\" class=\"wp-caption-text\"><em>Figure 31.4\u00a0<\/em>Gran Dolina site.\u00a0\u201c<a href=\"https:\/\/commons.wikimedia.org\/wiki\/File:Gran_Dolina_2012_(Sierra_de_Atapuerca).jpg\">Gran Dolina 2012 (Sierra de Atapuerca)<\/a>\u201d by Mario Modesto Mata is licensed under <a href=\"https:\/\/creativecommons.org\/licenses\/by-sa\/2.0\/deed.en\">CC BY-SA 2.0<\/a>.<\/p>\n<\/div>\n<h2><b>PHYSICAL CHARACTERISTICS<\/b><\/h2>\n<p>The 800 kya <i>H. antecessor<\/i> material was more derived than earlier <i>erectus<\/i> forms, possessing a larger brain (1,000\u20131,150 cc) and a more modern skull and face. They had low foreheads and possessed an <b>occipital bun<\/b> (a bun or chignon is the hairstyle wherein one winds long hair into a doughnut shape on the top or back of the head), as did neandertals (see neandertal features in Figure 31.5). It has been suggested that the purpose of that posterior protrusion is to balance the weight of the anterior portion of the skull and face. Based upon cranial anatomy, they are thought to have been capable of detecting the same range of sounds as modern humans and were possibly right-handed. They were almost as tall as modern humans at 5.5\u20136&#8242; (1.6\u20131.8 m), with males weighing ~200 lb (90\u00a0kg).<\/p>\n<div id=\"attachment_315\" style=\"width: 906px\" class=\"wp-caption aligncenter\"><img loading=\"lazy\" decoding=\"async\" aria-describedby=\"caption-attachment-315\" class=\"wp-image-315 size-full\" src=\"https:\/\/s3-us-west-2.amazonaws.com\/courses-images\/wp-content\/uploads\/sites\/2461\/2017\/09\/13202203\/image20-1.jpg\" alt=\"8.20\" width=\"896\" height=\"430\" \/><\/p>\n<p id=\"caption-attachment-315\" class=\"wp-caption-text\"><em>Figure 31.5\u00a0<\/em>Neandertal cranial characteristics.\u00a0\u201c<a href=\"https:\/\/commons.wikimedia.org\/wiki\/File:Neanderthal_cranial_anatomy.jpg\">Neanderthal cranial anatomy<\/a>\u201d by Jason Potter is licensed under <a href=\"https:\/\/creativecommons.org\/licenses\/by-sa\/2.5\/deed.en\">CC BY-SA 2.5<\/a>.<\/p>\n<\/div>\n<\/div>\n<div class=\"thirtyone\">\n<div class=\"textbox key-takeaways\">\n<h3>Review of Derived Characteristics<\/h3>\n<ul>\n<li>Increased encephalization.<\/li>\n<li>Occipital bun.<\/li>\n<li>Tall like Turkana Boy.<\/li>\n<\/ul>\n<\/div>\n<\/div>\n\n\t\t\t <section class=\"citations-section\" role=\"contentinfo\">\n\t\t\t <h3>Candela Citations<\/h3>\n\t\t\t\t\t <div>\n\t\t\t\t\t\t <div id=\"citation-list-216\">\n\t\t\t\t\t\t\t <div class=\"licensing\"><div class=\"license-attribution-dropdown-subheading\">CC licensed content, Shared previously<\/div><ul class=\"citation-list\"><li>The History of our Tribe: Hominini. <strong>Authored by<\/strong>: Barbara Welker. <strong>Provided by<\/strong>: SUNY Geneseo. <strong>Located at<\/strong>: <a target=\"_blank\" href=\"https:\/\/milnepublishing.geneseo.edu\/the-history-of-our-tribe-hominini\/\">https:\/\/milnepublishing.geneseo.edu\/the-history-of-our-tribe-hominini\/<\/a>. <strong>Project<\/strong>: Open SUNY Textbooks. <strong>License<\/strong>: <em><a target=\"_blank\" rel=\"license\" href=\"https:\/\/creativecommons.org\/licenses\/by-nc-sa\/4.0\/\">CC BY-NC-SA: Attribution-NonCommercial-ShareAlike<\/a><\/em><\/li><\/ul><\/div>\n\t\t\t\t\t\t <\/div>\n\t\t\t\t\t <\/div>\n\t\t\t <\/section>","protected":false},"author":62,"menu_order":17,"template":"","meta":{"_candela_citation":"[{\"type\":\"cc\",\"description\":\"The History of our Tribe: Hominini\",\"author\":\"Barbara Welker\",\"organization\":\"SUNY Geneseo\",\"url\":\"https:\/\/milnepublishing.geneseo.edu\/the-history-of-our-tribe-hominini\/\",\"project\":\"Open SUNY Textbooks\",\"license\":\"cc-by-nc-sa\",\"license_terms\":\"\"}]","CANDELA_OUTCOMES_GUID":"","pb_show_title":"on","pb_short_title":"","pb_subtitle":"","pb_authors":[],"pb_section_license":""},"chapter-type":[],"contributor":[],"license":[],"class_list":["post-216","chapter","type-chapter","status-publish","hentry"],"part":144,"_links":{"self":[{"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/pressbooks\/v2\/chapters\/216","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/pressbooks\/v2\/chapters"}],"about":[{"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/wp\/v2\/types\/chapter"}],"author":[{"embeddable":true,"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/wp\/v2\/users\/62"}],"version-history":[{"count":2,"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/pressbooks\/v2\/chapters\/216\/revisions"}],"predecessor-version":[{"id":350,"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/pressbooks\/v2\/chapters\/216\/revisions\/350"}],"part":[{"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/pressbooks\/v2\/parts\/144"}],"metadata":[{"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/pressbooks\/v2\/chapters\/216\/metadata\/"}],"wp:attachment":[{"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/wp\/v2\/media?parent=216"}],"wp:term":[{"taxonomy":"chapter-type","embeddable":true,"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/pressbooks\/v2\/chapter-type?post=216"},{"taxonomy":"contributor","embeddable":true,"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/wp\/v2\/contributor?post=216"},{"taxonomy":"license","embeddable":true,"href":"https:\/\/courses.lumenlearning.com\/suny-history-of-our-tribe\/wp-json\/wp\/v2\/license?post=216"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}